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Recombination in bacteria and in viruses that infect bacteria

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In E. coli, four Hfr strains donate the markers shown in the order given:

strain 1---Q---W---D---M---T
strain 2---A---X---P---T---M
strain 3---B---N---C---A---X
strain 4---B---Q---W---D---M

All these Hfr strains are derived from the same F+ strain.

What is the order of these markers on the circular chromosome of the original F+?

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The expert examines recombination in bacteria and in viruses that infect bacteria.

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